SJ Brown, TD Shippy, RE Denell, and RW Beeman (2004) Homeotic genes in Tribolium (Coleoptera) in The Encyclopedia of Entomology (Ed: Herb Oberlander).

RW Beeman, MD Lorenzen, Z Doyungan, TD Shippy, J Stuart, K Snow, J Savard, and SJ Brown (2004) Genetic linkage maps of the red flour beetle, Tribolium castaneum, based on bacterial artificial chromosomes and expressed sequence tags . Genetics in press.


A Burke and SJ Brown (2003) Homeotic Genes: Animals in Keywords in Evolution and Development (Ed: Brian K. Hall; University of Harvard Press) pp. 174-184.

SJ Brown, RE Denell, and RW Beeman (2003) Beetling around the genome . Genetical Research 82:155-161.

SJ Brown (2003) Entomological Contributions to Genetics: Studies on Insect Germ Cells Linked Genes to Chromosomes and Chromosomes to Mendelian Inheritance . Archives of Insect Biochemistry and Physiology 53:115-118.

MD Lorenzen, AJ Berghammer, SJ Brown, RE Denell, M Klingler, RW Beeman (2003) piggyBac-mediated germline transformation in the beetle Tribolium castaneum . Insect Molecular Biology 5:433-40.


SJ Brown, J Fellers, TD Shippy, EA Richardson, M Maxwell, J Stuart, and RE Denell (2002) Sequence of the ANTC portion of the Tribolium Homeotic Complex . Genetics 160:1064-74.

MD Lorenzen, SJ Brown, RE Denell, and RW Beeman (2002) Cloning and Characterization of the tryptophan oxygenase and kynurenine 3-monooxygenase genes from Tribolium castaneum . Genetics 160:225-234.

SJ Brown, TD Shippy, RW Beeman, and RE Denell (2002) Tribolium Hox genes repress antennal development in the gnathos and trunk . Molecular Phylogenetics and Evolution 24:384-387.

MD Lorenzen, SJ Brown, RE Denell, and RW Beeman (2002) Transgene expression from the Tribolium castaneum Polyubiquitin promoter . Insect Molecular Biology 11:399-407.


MA DeCamillis, DL Lewis, SJ Brown, RW Beeman, and RE Denell (2001) Interactions of the Tribolium Sex combs reduced and proboscipedia orthologs in embryonic labial development . Genetics 159:1643-1648.

W Nie, B Stronach, G Panganiban, TD Shippy, SJ Brown, and RE Denell (2001) Molecular characterization of Tclabial and the 3' end of the Tribolium Homeotic Complex . Development Genes and Evolution 211:244-251.

MS Haas, SJ Brown, and RW Beeman (2001) Homeotic evidence for the appendicular origin of the labrum in Tribolium castaneum . Development Genes and Evolution 211:96-102.

MS Haas, SJ Brown, and RW Beeman (2001) Pondering the Procephalon: segmental origin of the labrum . Development Genes and Evolution 211:89-95.

C Curtis, JA Brisson, MA DeCamillis, TD Shippy, SJ Brown, and RE Denell (2001) Molecular characterization of Cephalothorax, the Tribolium ortholog of Sex combs reduced . Genesis 30:12-20.

SJ Brown, J Fellers, TD Shippy, RE Denell, M Stauber, and U Schmidt-Ott (2001) A strategy for mapping bicoid on the phylogenetic tree . Current Biology 11:R43-44.


SJ Brown, M DeCamillis, K Gonzales, M Denell, S Haas, RW Beeman, W Nie, and RE Denell (2000) Implications of the Tribolium Deformed mutant phenotype for the evolution of Hox gene function . PNAS 97:4510-4515.

TD Shippy, SJ Brown, RE Denell (2000) maxillopedia is the Tribolium ortholog of proboscipedia . Evolution and Development 2:145-151.

TD Shippy, J Guo, SJ Brown, RW Beeman, MS Haas, MD Lorenzen, and RE Denell (2000) Analysis of the maxillopedia expression pattern and larval cuticular phenotype in wild-type and mutant Tribolium . Genetics 155:721-731.


R Bennett, SJ Brown, and RE Denell (1999) Molecular and Genetic Analysis of the Tribolium Ultrabithorax ortholog, Ultrathorax . Development Genes and Evolution 209:608-619.

RW Beeman and SJ Brown (1999) RAPD-based genetic linkage maps of Tribolium castaneum . Genetics 153:333-338.

SJ Brown, JP Mahaffey, M Lorenzen, RE Denell, and JW Mahaffey (1999) Using RNAi to investigate orthologous homeotic gene function during development of distantly related insects . Evolution and Development 1:11-15.

SJ Brown, S Holtzman, TC Kaufman, and RE Denell (1999) Characterization of the Tribolium Deformed ortholog and its ability to directly regulate Deformed target genes in the rescue of a Drosophila Deformed null mutant . Development Genes and Evolution 209:389-398.


TD Shippy, SJ Brown, and RE Denell (1998) Molecular characterization of the Tribolium abdominal-A ortholog and implications for the products of the Drosophila gene . Development Genes and Evolution 207:446-452.


A Guichet, JWR Copeland, M Erdélyi, D Hlousek, P Závorszky, J Ho, SJ Brown, A Percival-Smith, HM Krause, and A Ephrussi (1997) The nuclear receptor homologue Ftz-F1 and the homeodomain protein Ftz are mutually dependent cofactors . Nature 385:548-552.

SJ Brown, JK Parrish, RW Beeman, and RE Denell (1997) Molecular characterization and embryonic expression of the even-skipped ortholog of Tribolium castaneum . Mech. Dev. 61:165-173.


JJ Stuart, MJ De Gortari, PS Hall, ME Maxwell, G Mocelin, SJ Brown, and WM Muir (1996) Useful DNA polymorphisms are identified by snapback, a midrepetitive element in Tribolium castaneum . Genome 39:568-578.

J Sanchez-Salazar, MT Pletcher, RL Bennett, SJ Brown, TJ Dandamudi, RE Denell, and JS Doctor (1996) The Tribolium decapentaplegic gene is similar in sequence, structure, and expression to the Drosophila dpp gene . Roux's Arch. Dev. Biol. 206:237-246.

YB Li, SJ Brown, B Hausdorf, D Tautz, RE Denell, and R Finkelstein (1996) Two orthodenticle-related genes in the short-germ beetle Tribolium castaneum . Roux's Arch. Dev. Biol. 206:35-45.

F Falciani, B Hausdorf, R Schröder, M Akam, D Tautz, RE Denell, and SJ Brown (1996) Class 3 Hox genes in insects and the origin of zen . PNAS USA 93:8479-8484.

RE Denell, SJ Brown, and RW Beeman (1996) Evolution of the organization and function of insect homeotic complexes . Sem. Cell Dev. Biol. 7:527-538.

RW Beeman, MS Thomson, JM Clark, MA DeCamillis, SJ Brown, and RE Denell (1996) Woot, an active gypsy-class retrotransposon in the flour beetle, Tribolium castaneum, is associated with a recent mutation . Genetics 143:417-426.

SJ Brown and RE Denell (1996) Segmentation and dorsal-ventral patterning in Tribolium . Sem. Cell Dev. Biol. 7:553-560.


SJ Brown, NH Patel, and RE Denell (1994) Embryonic expression of the single Tribolium engrailed homolog . Development Genetics 15:7-18 (cover article).

SJ Brown, JK Parrish, and RE Denell (1994) Genetic control of early embryogenesis in the red flour beetle Tribolium castaneum . American Zoologist 34:343-352.

SJ Brown, RB Hilgenfeld, and RE Denell (1994) The beetle Tribolium castaneum has a fushi tarazu homolog expressed in stripes during segmentation . PNAS 91:12922-12926.


JJ Stuart, SJ Brown, RW Beeman, and RE Denell (1993) The Tribolium homeotic gene Abdominal is homologous to abdominal-A of the Drosophila bithorax complex . Development 117:233-243.

RW Beeman, SJ Brown, JJ Stuart, and RE Denell (1993) The evolution of genes regulating developmental commitments in insects and other animals in Evolutionary Conservation of Developmental Mechanisms (Ed: A.C. Spradling; New York: Wiley-Liss, Inc.) pp. 71-84.

RW Beeman, JJ Stuart, SJ Brown, and RE Denell (1993) Structure and function of the Homeotic gene complex (HOM-C) in the beetle Tribolium castaneum . BioEssays 15: 439-444 (cover article).


JJ Stuart, SJ Brown, RW Beeman, and RE Denell (1991) A deficiency of the homeotic complex of the beetle Tribolium castaneum . Nature 350:72-74.


RW Beeman, SJ Brown, JJ Stuart, and RE Denell (1990) Homeotic genes of the red flour beetle in Molecular Insect Science (Eds: H. Hagedorn, J. Hildebrand, M. Kidwell, and J. Law; New York: Plenum Press) pp. 21-29.

SJ Brown, JK Henry, WC Black, and RE Denell (1990) Molecular genetic manipulation of the red flour beetle: Genome organization and cloning of a ribosomal protein gene . Insect Biochem 20:185-193.