Kansas State University


Konza Prairie Biological Station


Long Term Ecological Research



Anthony Joern


2000-Present         1990-1999         1980-1989         1975-1979     



Behmer, S.T. and A. Joern. 2008. Coexisting generalist herbivores occupy unique nutritional feeding niches.  Proceedings of the National Academy of Sciences, USA 105: 1977-1982.

Jonas, J.L. and A. Joern. 2007. Grasshopper (Orthoptera: Acrididae) communities respond to fire, bison grazing and weather in North American tallgrass prairie: a long-term study. Oecologia 153: 699-711.

Jonas, J.L., G.W.T. Wilson, P.M. White, and A. Joern.  2007.  Consumption of mycorrhizal and saprophytic fungi by Collembola in grassland soils.  Soil Biology and Biochemistry 39: 2594-2602.

Sword, G.A. Sword, L. B. Senior, J. F. Gaskin, and A. Joern. 2007. Double trouble for grasshopper molecular systematics: Intra-individual heterogeneity of both mitochondrial 12S-valine-16S and nuclear internal transcribed spacer ribosomal DNA sequences in Hesperotettix viridis (Orthoptera: Acrididae).  Systematic Entomology 32: 420-428.

Logan, J.D., Wolesensky, W. and A. Joern.  2007.  Insect Development Under Predation Risk, Variable Temperature, and Variable Food Quality.  Mathematical Biosciences and Engineering 4: 47-65.

Branson, D.H., A. Joern, and G.A. Sword. 2006.  Sustainable Management of Insect Herbivores in Grassland Ecosystems: New Perspectives in Grasshopper Control. BioScience 56: 743-755.

Joern, A., B.J. Danner, J.D. Logan, and W. Wolesensky.  2006.  Natural History of Mass-action in Predator-prey Models: A Case Study from Wolf Spiders and Grasshoppers.  American Midland Naturalist 156: 52-62.

Logan, J.D., W. Wolesensky and A. Joern.  2006.  Temperature-Dependent Phenology and Predation in Arthropod Systems.  Ecological Modelling 196: 471-482.

Joern, A. 2005.  Disturbance by Fire and Bison Grazing Modulate Grasshopper Assemblages In Tallgrass Prairie. Ecology 86:861-873.

Joern, A. and S. Mole.  2005.  The Plant Stress Hypothesis and Variable Responses by Blue Grama Grass (Bouteloua gracilis) To Water, Mineral Nitrogen, and Insect Herbivory. Journal of Chemical Ecology 31: 2069-2090.

Sword, G.A., A. Joern and L.B. Senior.  2005.  Host plant associated genetic differentiation in the snakeweed grasshopper, Hesperotettix viridis (Orthoptera: Acrididae). Molecular Ecology 14:  2197-2205.

Joern, A., J.D. Logan, and W. Wolesensky.  2005. Effect of global climate change on agricultural pests: possible impacts and dynamics at population, species-interaction and community levels.  Pages 321-362 IN  R. Lal, N. Uphoff, B.A. Stewart and D.O. Hansen (eds.).  Climate Change and Global Food Security.  CRC Press, Boca Raton, FL.

Wolesensky, W., A. Joern, & J. D. Logan, 2005. A model of digestion modulation in grasshoppers. Ecological Modelling, 188, 358-373.

Joern, A. 2004. Variation in Grasshopper Densities in Response to Fire Frequency and Bison Grazing in Tallgrass Prairie.  Environmental Entomology 33: 1617-1625.

Joern, A.  Grasshoppers.  2004. Encyclopedia of the Great Plains.  D.J. Wishart (ed).  University of Nebraska Press, Lincoln, NE.

Stoner, K.L. and A. Joern.  2004.  Landscape vs. Local Habitat Scale Influences to Insect Communities from Tallgrass Prairie Remnants. Ecological Applications 14: 1306-1320.

Ledder, G., J.D. Logan and A. Joern.  2004.  Dynamic energy budget models with size-dependent hazard rates.  Bulletin of Mathematical Biology 48: 605-622.

Botkin, D. and others 2004.  Ten suggestions to strengthen the science of ecology.  BioScience 54: 345-351.

Danner, B.J. and A. Joern.  2004.  Development, growth and egg production of Ageneotettix deorum (Orthoptera: Acrididae) in response to spider predation risk and elevated resource quality.  Ecological Entomology 29:1-11

Logan, J.D., A. Joern, and W. Wolesensky.  2004.  Control of CNP homeostasis in herbivore consumers through differential assimilation. Bulletin of Mathematical Biology 66: 707-725.

Logan, J.D., A. Joern and W. Wolesensky.  2003.  Mathematical model of consumer homeostasis control in plant-herbivore dynamics. Mathematical and Computer Modelling 40: 447-456.

Danner, B.J. and A. Joern.  2003.  Resource mediated impact of spider predation risk on performance in the grasshopper Ageneotettix deorum (Orthoptera: Acrididae).  Oecologia 137: 352-359.

Danner, B.J. and A. Joern. 2003.  Stage-Specific Behavioral Responses of Ageneotettix deorum (Orthoptera: Acrididae) in the Presence of Lycosid Spider Predators.  Journal of Insect Behavior 16: 453-464.

Logan, J.D., A. Joern, & W. Wolesensky.  2003.  Chemical reactor models of optimal digestion efficiency with constant foraging costs. Ecological Modelling 168: 25-38.

Joern, A. 2002.  Context-dependent foraging and enemy-free space: Grasshopper sparrows (Ammodramus savannarum) searching for grasshoppers (Acrididae).  Ecoscience 9: 231-240.

Logan, J.D., A. Joern and W. Wolesensky.  2002.  Location, Time and Temperature Dependence of Digestion in Simple Animal Tracts.  Journal of Theoretical Biology 216: 5-18.

Winemiller, K.O., E.R. Pianka, L.J. Vitt, and A. Joern.  2001. Food Web Laws or Niche Theory?  Six Independent Empirical Tests.  American Naturalist 158:  193-199.

Joern, A. 2000.  What are the consequences of non-linear ecological interactions for grasshopper control strategies?  pp. 131-144, IN  J.A. Lockwood, A.V. Latchininsky and M. Sergeev (eds.). Grasshoppers and Grassland Health.  Kluwer Academic Publishers.  Dordrecht, The Netherlands.

Holz, J.C., K.D. Hoagland, and A. Joern.  2000.  Aquatic food web interactions:  micrososms as lake models. Pp. 305-324  In Tested Studies for Laboratory Teaching, Volume 21 (S.J. Karcher, editor).  Proceedings of the 21st Workshop/Conference of the Association for Biological Laboratory Instructors (ABLE).

Oedekoven, M.A. and A. Joern. 2000.  Plant Quality and Spider Predation Affects Grasshoppers (Acrididae):  Food-Quality-Dependent Compensatory Mortality.  Ecology 81: 66-77.

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Traxler, M. and A. Joern.  1999.  Performance tradeoffs for two hosts within and between populations of the oligophagous grasshopper Hesperotettix viridis (Acrididae).  Oikos 87: 239-250.

Oedekoven, M.A. and A. Joern.  1998.  Stage-based mortality of grassland grasshoppers (Acrididae) from wandering spider (Lycosidae) predation.  Acta Oecologia 19: 507-515.

Joern, A. and S.T. Behmer.  1998. Impact of diet quality on demographic attributes in adult grasshoppers and the nitrogen limitation hypothesis. Ecological Entomology 23: 174-184.

Appel, H.M. and A. Joern.  1998.  Gut physicochemistry of grassland grasshoppers.  Journal of Insect Physiology 44:  693-700.

Joern, A. and S.T. Behmer.  1997.  Importance of dietary nitrogen and carbohydrates to survival, growth and reproduction in adult Ageneotettix deorum (Orthoptera: Acrididae).  Oecologia 112:  201-208.

Fuller, R.C. and A. Joern.  1996.  Grasshopper susceptibility to predation in response to vegetation cover, prey diversity and patch size.  Journal of Orthoptera Research 5: 175-183.

Joern, A. and K.D. Hoagland. 1996. In defense of whole-community bioassays for risk assessment.  Environmental Toxicology & Chemistry 15:  407-409.

Arenz, C.L. and A. Joern.  1996.  Prairie legacies - invertebrates.  Pp. 91- 109 In Prairie Conservation: Preserving North America’s Most Endangered Ecosystem (F.B. Samson and FL Knopf, eds.).  Island Press, Covello, CA.

Belovsky, G.E. and A. Joern.  1995.   The dominance of different regulating mechanisms for rangeland grasshoppers.  Pp. 359-386  in N. Cappuccino and P.W. Price (eds).  Population Dynamics:  New Approaches and Synthesis.  Academic Press, San Diego.

Joern, A. and K.H. Keeler (eds).  1995.  The Changing Prairie:  North American Grasslands.  Oxford University Press.  NY

Joern, A. 1995.  The entangled bank:  species interactions and grassland dynamics.  Chapter 6 in A. Joern and K.H. Keeler (eds).  The Changing Prairie: North American Grasslands.  Oxford University Press. Oxford. NY

Joern, A. and K.H. Keeler. 1995.  Getting the lay of the land:  introduction to North American grasslands.  Chapter 1 in A. Joern and K.H. Keeler (eds).  The Changing Prairie:  North American Grasslands. Oxford University Press. NY

Mole, S.  and A. Joern.  1994.  Feeding Behavior of Graminivorous Grasshoppers in Response to Host-plant Extracts, Alkaloids and Tannins.  Journal Chemical Ecology 20:  3097-3109.

Behmer, S.T. and A. Joern.  1994.  The influence of proline on diet selection:  sex-specific feeding preferences by the grasshoppers Ageneotettix deorum and Phoetaliotes nebrascensis (Orthoptera: Acrididae).  Oecologia 98: 76-82.

Mole, S., A. Joern, M.H. O'Leary and S. Madhavan.  1994.  Spatial and temporal variation in carbon isotope discrimination in prairie graminoids.  Oecologia 97: 316-321.

Yang, Y. and A. Joern. 1994.  Gut size changes in response to variable food quality and body size in grasshoppers.  Functional Ecology 8: 36-45.

Yang, Y. and A. Joern. 1994.  Compensatory feeding in response to variable food quality by Melanoplus differentialis.  Physiological Entomology  19:  75-82.

Yang, Y. and A. Joern.  1994.  Influence of Diet Quality, Developmental Stage, and Temperature on Food Residence Time in the Grasshopper Melanoplus differentialis.  Physiological Zoology 67: 598-616.

Alward, R.D. and A. Joern. 1993.  Plasticity and overcompensation in grass responses to herbivory.  Oecologia 95:  358-364.

Joern, A. and G. Klucas. 1993. Intra- and Interspecific Competition in Adults of Two Abundant Grasshopper (Orthoptera: Acrididae) from a Sandhills Grassland.  Environmental Entomology 22: 352-361.

Kaspari, M.E. and A. Joern.  1993.  Prey choice by three insectivorous grassland birds: reevaluating opportunism.  Oikos 68:  414-430.

Mole, S. and A. Joern.  1993.  Foliar Phenolics of Nebraska Sandhills Prairie Graminoids:  Between-years, Seasonal and Interspecific Variation.  Journal Chemical Ecology 19: 1861-1874.

Behmer, S.T. and A. Joern.  1993.  Diet choice by a grass-feeding grasssshopper based on the need for a limiting nutrient.  Functional Ecology 7:  522-527.

Berry, J.S., G. Belovsky, A. Joern, W.P. Kemp, and J. Onsager.  1993.  Object-oriented Simulation Model of Rangeland Grasshopper Population Dynamics.  Proceedings Fourth Conference on AI Simulation and Planning in High-Autonomy Systems.  IEEE Computer Society Press.  

Joern, A. 1992.  Variable Impact of Avian Predation on Grasshopper Assemblies in Sandhills Grassland.  Oikos 64: 458-463.

Joern, A. and S.B. Gaines. 1990. Population dynamics and regulation in grasshoppers. pp. 415-482 In R.F. Chapman and A. Joern (eds.). Biology of Grasshoppers. John Wiley and Sons. NY

Chapman, R.F. and A. Joern (eds.). 1990. Biology of Grasshoppers. John Wiley & Sons. NY.

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Joern, A. 1989. Insect herbivory in the transition to California annual grasslands: did grasshoppers deliver the coup de grass? pp. 117-134 In L. Huenneke and H. Mooney (eds). Grassland Structure and Function: California Annual Grasslands.  Kluwer Academic Publishers.  Dordrecht, The Netherlands.

Joern, A. 1988. Foraging behavior and switching by the grasshopper sparrow, Ammodramus savannarum, searching for multiple prey in a heterogeneous environment.  American Midland Naturalist 119: 225-234.

Joern, A. and R. Alward. 1988. Effects of nitrogen fertilization on choice among grasses by the grasshopper Phoetaliotes nebrascensis (Orthoptera: Acrididae).  Annals Entomological Society America 81: 240-244.

Heidorn, T.J. and A. Joern. 1987. Feeding preference and spatial distribution of grasshoppers  (Acrididae) in response to nitrogen fertilization of Calamovilfa longifolia. Functional Ecology 1: 369-375.

Joern, A. 1987. Behavioral responses underlying ecological patterns: resource use in grasshoppers. pp. 137-161 In J.C. Capinera (Ed). Rangeland Pest Management, A Shortgrass Prairie Perspective. Westview Press. Boulder, CO.

Joern, A., R. Mitschler and H. O'Leary. 1986. Activity and time budgets in common sandhills prairie grasshoppers. Journal Kansas Entomological Society 59: 1-6.

Joern, A. 1986. Experimental study of avian predation on coexisting grasshopper populations (Orthoptera: Acrididae) in a sandhills grassland.  Oikos 46: 243-249.

Joern, A. 1986. Resource partitioning by grasshopper species from grassland communities. pp. 75-100. In D.A. Nickle (ed.). Proceedings IV Triennial Meeting Pan American Acridological Society. PAAS.

Joern, A. and K.P. Pruess. 1986. Temporal constancy in grasshopper assemblies (Orthoptera: Acrididae). Ecological Entomology 11: 379-385.

Joern, A. 1985. Expanding our horizons in grasshopper ecology. pp. 93-96. In D.A. Nickle (ed.). 3rd Triennial Proceedings Pan Amer. Acridological Society. Edward Bros. Ann Arbor, Michigan

Joern, A. 1985. Grasshopper dietary (Orthoptera: Acrididae) from a Nebraska sandhills prairie. Transaction Nebraska Academy Science 8:21-32.

Heidorn, T. and A. Joern. 1984. Differential herbivory on C3 versus C4 grasses by the grasshopper Ageneotettix deorum (Orthoptera: Acrididae). Oecologia 65:19-25.

Joern, A. 1983. Small-scale Displacements of Grasshoppers (Orthoptera: Acrididae) Within Arid Grasslands. Journal Kansas Entomological Society 56:131-139.

Joern, A. 1983. Host Plant Utilization by Grasshoppers (Orthoptera: Acrididae) from a Sandhills Prairie. Journal Range Management 36:793-797.

Joern, A. 1982. Vegetation structure and microhabitat selection in grasshoppers (Orthoptera: Acrididae). Southwestern Naturalist 27:197-209.

Joern, A. 1982. Distributions, Densities and Relative Abundances of Grasshoppers (Orthoptera: Acrididae) in a Nebraska Sandhills Prairie.  Prairie Naturalist 14:37-45.

Joern, A. and N. Rudd. 1982. Impact of Predation by the Robber Fly Proctacanthus milbertii (Diptera: asilidae) on Grasshopper (Orthoptera: acrididae) Populations. Oecologia  55:42-46.

Joern, A.  1981.  Importance of behavior and coloration in the control of body temperature by Brachystola magna Girard (Orthoptera: Acrididae). Acrida 10: 117-130.

Joern, A. and L.R. Lawlor. 1981. Guild structure in grasshopper assemblages based on food and microhabitat resources. Oikos 37:93-104.

Joern, A. 1981.  Expanding Our horizons in grasshopper ecology.  pp. 93-95 in D.A. Nickle (Ed). Proceedings 3rd Triennial Meeting, Pan American Acridological Society.

Joern, A. and L.R. Lawlor. 1980. Food and Microhabitat Utilization by Grasshoppers from Arid Grasslands: Comparisons with Neutral Models. Ecology  61:591-599.

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Joern, A. 1979. Feeding Patterns in Grasshoppers (Orthoptera: Acrididae): Factors Influencing Diet Specialization. Oecologia  38:325-347.

Joern, A. 1979. Resource utilization and community structure in assemblages of arid grassland grasshoppers (Orthoptera: Acrididae). Transaction American  Entomological Society 150:253-300.

Joern, A. 1979. Resource utilization by grasshopper assemblages:  Some Implications Concerning Interactions Among Species.  pp. 112 in M. Tyrkus, I.J. Cantrall & C.S. Carbonell (eds). Proc. 2nd Triennial Meetings.  Pan American Acridological Society.

Fry, B., A. Joern, and P.L. Parker. 1978. Grasshopper food web analysis: use of carbon isotope rates to examine feeding relationships among terrestrial herbivores. Ecology 59:498-506.

Joern, A. 1977. Guild and Community Structure in Primary Consumers: Resource Utilization in Arid Grassland Communities. Ph.D. Thesis. University of Texas at Austin. 325 p.

Otte, D. and A. Joern. 1977. On feeding patterns in desert grasshoppers and the evolution of specialized diets. Transactions Philadelphia Academy Sciences 128:89-126

Joern, A. 1976. Grasshopper affinities and habitat relationships in the Solitario. pp. 121-124. In D. Kinard (ed.), The Solitario: A Natural History Survey. No. 9 LBJ School of Public Affairs. Austin, Texas.

Otte, D. and A. Joern. 1975. Insect territoriality and its evolution:  population studies of desert grasshoppers on creosote bushes.  Journal Animal Ecology 44:29-54.

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