Laboratory for Landscape and Conservation Ecology

at Kansas State University







bullet Lab Director
bullet Lab Personnel









Sinauer Associates
Coming soon!


Essentials of Landscape Ecology
Kimberly A. With

Essentials of Landscape Ecology
investigates the causes and consequences of environmental heterogeneity—the reciprocal effects of pattern and process—ranging from human land use and management considerations to the ecological and evolutionary considerations of biota within landscapes, including aquatic and marine landscapes.

In this new, fully up-to-date and comprehensive text, the principles and applications of landscape ecology are presented in an engaging and accessible format. Suitable for upper-level undergraduate or graduate courses in landscape ecology, this textbook could also be used as a supplemental text in courses on forestry, resource and wildlife management, conservation and restoration biology, and land-use planning. For the professional, this book provides a complete reference to the important concepts, methods, and applications of landscape ecology. This book is thus relevant to ecologists, resource management professionals, wildlife and fisheries biologists, conservation biologists, land-use planners, and policy makers who are interested in understanding the consequences of spatial heterogeneity and/or are charged with incorporating landscape ecological principles into their research or management programs.



1. What is Landscape Ecology? Foundation and Core Concepts
2. Scaling Issues in Landscape Ecology
3. Landscape Heterogeneity and Dynamics
4. Landscape Pattern Analysis
5. Landscape Connectivity
6. Landscape Effects on Individual Movement and Dispersal: Behavioral Landscape Ecology
7. Landscape Effects on Population Distributions and Dynamics
8. Landscape Effects on Population Spatial Spread: Range Shifts, Invasive Spread, and Landscape Epidemiology
9. Landscape Effects on Gene Flow and Population Genetic Structure: Landscape Genetics
10. Landscape Effects on Community Structure and Dynamics
11. Landscape Effects on Ecosystem Processes and Function


9. With, K. A.  2007.  Invoking the ghosts of landscapes past to understand the landscape ecology of the present...and the future.  Pp. 43-58 in Temporal Explicitness in Landscape Ecology:  Wildlife Responses to Changes in Time (J. A. Bissonette and I. Storch, editors).  Springer, New York.
8. Taylor, P., L. Fahrig, and K. A. With.  2006.  Landscape connectivity: back to the basics. Pp. 29-43 in Connectivity Conservation (K. Crooks and M. Sanjayan, editors). Cambridge University Press, Cambridge, UK.
7. With, K. A.  2005.  Case study:  Assessing extinction risk in migratory songbirds:  the need for landscape-based demographic models.  Pp. 449-453 in Principles of Conservation Biology, 3rd edition (M. J. Groom, G. K. Meffe, C. R. Carroll, and contributors).  Sinauer, Sunderland, MA.
6. With, K. A.  2005.  Landscape conservation:  a new paradigm for the conservation of biodiversity.  Pp. 238-247 in Issues and Perspectives in Landscape Ecology (J. A. Wiens and M. R. Moss, editors).  Cambridge University Press, Cambridge, UK.
5. With, K. A.  2004.  Metapopulation dynamics:  perspectives from landscape ecology.  Pp. 23-44 in Metapopulation Dynamics:  Ecology, Genetics, and Evolution of Metapopulations (I. Hanski and O. E. Gaggiotti, editors).  Academic Press, San Diego, CA.     [Request reprint]
4. With, K. A. 2002.  Using percolation theory to assess landscape connectivity and effects of habitat fragmentation. Pp. 105-130 in Applying Landscape Ecology in Biological Conservation (K. J. Gutzwiller, editor). Springer-Verlag, New York.
3.  With, K. A. 2002. Landscape connectivity and metapopulation dynamics. Pp. 208-227 in Learning Landscape Ecology: A Practical Guide to Concepts and Techniques (S.E. Gergel and M. G. Turner, editors). Springer-Verlag, New York.
2. With, K. A. 1999. Is landscape connectivity necessary and sufficient for wildlife management? Pp. 97-115 In Forest Fragmentation: Wildlife and Management Implications (J. A. Rochelle, L. A. Lehmann, and J. Wisniewski, editors). Brill Academic Publishers, Leiden, The Netherlands.  
1. With, K. A. 1994. Updated 2010.  McCown's Longspur (Calcarius mccownii). In The Birds of North America, No. 96 (A. Poole and F. Gill, eds.). Philadelphia: The Academy of Natural Sciences; Washington, DC.: The American Ornithologists' Union.   Available at: The Birds of North America Online


55.  Mendes, P., K. A. With, L. Signorelli, and P. De Marco Jr. 2017. The relative importance of local versus landscape variables on site occupancy in bats of the Brazilian Cerrado. Landscape Ecology 32: 745-762.
54.  Signorelli, L., R. P. Bastos, P. De Marco Jr., and K. A. With.  2016.  Landscape context affects site occupancy of pond-breeding anurans across a disturbance gradient in the Brazilian Cerrado. Landscape Ecology 31: 1997-2012.
53.  With, K. A.  2016.  Are landscapes more than the sum of their patches?  Landscape Ecology 31:  969-980.
52.  With, K. A.  2015.  How fast do migratory songbirds have to adapt to keep pace with rapidly changing landscapes?  Landscape Ecology 30:  1351-1361.
51.  Skelsey, P., K. A. With, and K. A. Garrett.  2013.  Pest and disease management:  Why we shouldn't go against the grainPLoS ONE 8(9): e75892.
50.  Skelsey, P., K. A. With, and K. A. Garrett.  2013.  Why dispersal should be maximized at intermediate scales of heterogeneityTheoretical Ecology 6: 203-211.
49.  With, K. A., and D. M. Pavuk.  2012. Direct versus indirect effects of habitat fragmentation on community patterns in experimental landscapes. Oecologia 170: 517-528.
48.  Klug, P. E., J. Fill, and K. A. With. 2011. Spatial ecology of eastern yellowbelly racer (Coluber constrictor flaviventris) and Great Plains ratsnake (Pantherophis emoryi) in a contiguous tallgrass-prairie landscape. Herpetologica 67: 428-439.
47.  Blevins, E., S. M. Wisely, and K. A. With.  2011.  Historical processes and landscape context influence genetic structure in peripheral populations of the collared lizard (Crotaphytus collaris)Landscape Ecology 26: 1125-1136.
46.  With, K. A., and D. M. Pavuk.  2011.  Habitat area trumps fragmentation effects on arthropods in an experimental landscape systemLandscape Ecology 26: 1035-1048.
45.  Blevins, E., and K. A. With. 2011.  Landscape context matters: local vs. landscape effects on abundance and patch occupancy of collared lizards in managed grasslandsLandscape Ecology 26: 837-850.
44.  Klug, P. E., S. M. Wisely, and K. A. With. 2011. Population genetic structure and landscape connectivity of the Eastern Yellowbelly Racer (Coluber constrictor flaviventris) in the tallgrass prairie of northeastern Kansas, USA. Landscape Ecology 26: 281-294.
43.  Romero, S., J. F. Campbell, J. R. Nechols, and K. A. With. 2010. Search behavior of red flour beetle: response to habitat cues and patch boundariesEnvironmental Entomology 39: 919-929.
42.  Klug, P. E., S. L. Jackrel, and K. A. With.  2010.  Linking snake habitat use to nest predation risk in grassland birds: the dangers of shrub coverOecologia 162: 803-813.
41.  Klug, P. E., K. A. With, and S. M. Wisely. 2009. Microsatellite isolation and development in the Eastern yellowbelly racer (Coluber constrictor flaviventris) for use in landscape genetic studies. (Molecular Ecology Resources database accessions 37758–37769). Cited as: Molecular Ecology Resources Primer Development Consortium et al. 2009. Permanent Genetic Resources added to Molecular Ecology Resources Database 1 May 2009–31 July 2009. Molecular Ecology Resources 9: 1460-1466.
40.  Long, A. M., W. E. Jensen, and K. A. With. 2009. Orientation of Grasshopper Sparrow and Eastern Meadowlark nests in relation to wind direction. Condor 111: 395-399.
39.  Margosian, M. L. K. A. Garrett, J. M. S. Hutchinson, and K. A. With. 2009. Connectivity of the American agricultural landscape: Assessing the national risk of disease and crop pest spread. BioScience 59: 141-151.
38.  Romero, S., J. F. Campbell, J. R. Nechols, and K. A. With.  2009.  Movement behavior in response to landscape structure: role of functional grain.  Landscape Ecology 24: 39-51.
37.  Rahmig, C. J., W. E. Jensen, and K. A. With.  2009.  Grassland bird responses to land management in the largest remaining tallgrass prairieConservation Biology 23: 420-432
36.  With, K. A., A. W. King, and W. E. Jensen.  2008.  Remaining large grasslands may not be sufficient to prevent grassland bird declinesBiological Conservation 141: 3152-3167.
35.  Frey, C. M., W. E. Jensen, and K. A. With. 2008. Topographic patterns of nest placement and habitat quality for grassland birds in the Flint Hills.  American Midland Naturalist 160: 220-234.
34.  Scheiman, D. M., J. B. Dunning, Jr., and K. A. With. 2007. Metapopulation dynamics of Bobolinks occupying agricultural grasslands in the Midwestern United States.  American Midland Naturalist 158: 415-423.
33.  With, K. A., G. R. Schrott, and A. W. King.  2006. The implications of metalandscape connectivity for population viability in migratory songbirdsLandscape Ecology 21: 157-167.
32.  Schrott, G. R., K. A. With, and A. W. King.  2005.  Demographic limitations of the ability of habitat restoration to rescue declining populations Conservation Biology 191181-1193.  
31.  Schrott, G. R., K. A. With, and A. W. King.  2005.  On the importance of landscape history for assessing extinction riskEcological Applications 15:  493-506.
30.  With, K. A., and A. W. King.  2004.  The effect of landscape structure on community self-organization and critical biodiversityEcological Modelling 179:  349-366.
29.  With, K. A.  2004.  Assessing the risk of invasive spread in fragmented landscapesRisk Analysis 24: 803-815.  "Recommended" by Faculty of 1000 Biology
28.  With, K. A.  2002.  The landscape ecology of invasive spreadConservation Biology 16:  1192-1203.
27.  With, K. A., D. M. Pavuk, J. L. Worchuck, R. K. Oates, and J. L. Fisher.  2002.  Threshold effects of landscape structure on biological control in agroecosystemsEcological Applications 12: 52-65. 
26.  King, A. W., and K. A. With.  2002.  Dispersal success on spatially structured landscapes:  when do spatial pattern and dispersal behavior really matter?  Ecological Modelling 147: 23-39.
25.  Sakai, A. K., F. W. Allendorf, J. S. Holt, D. M. Lodge, J. Molofsky, K. A. With, S. Baughman, R. J. Cabin., J. E. Cohen, N. C. Ellstrand, D. E. McCauley, P. O'Neil, I. M. Parker, J. N. Thompson, and S. G. Weller.  2001.  The population biology of invasive speciesAnnual Review of Ecology and Systematics 32305-332.
24.  With, K. A., and A. W. King. 2001. Analysis of landscape sources and sinks: the effect of spatial pattern on avian demography. Biological Conservation 100: 75-88.  Recipient of the 2002 Award for Best Paper published in the field of landscape ecology (US-IALE)
23.  With, K. A., S. J. Cadaret, and C. Davis. 1999. Movement responses to patch structure in experimental fractal landscapes. Ecology 80: 1340-1353.  
22.  With, K. A., and A. W. King. 1999. Extinction thresholds for species in fractal landscapes. Conservation Biology 13: 314-326.  
21.  With, K. A., and A. W. King. 1999. Dispersal success in fractal landscapes: a consequence of lacunarity thresholds. Landscape Ecology 14: 73-82.
20.  With, K. A. 1997. The theory of conservation biology. Conservation Biology 11: 1436- 1440.
19.  With, K. A. 1997. The application of neutral landscape models in conservation biology. Conservation Biology 11: 1069-1080.
18.  With, K. A., and A. W. King. 1997. The use and misuse of neutral landscape models in ecology. Oikos 79: 219-229.
17.  With, K. A., R. H. Gardner, and M. G. Turner. 1997. Landscape connectivity and population distributions in heterogeneous environments. Oikos 78: 151-169.
16.  With, K. A., and T. O. Crist. 1996. Translating across scales: simulating species distributions as the aggregate response of individuals to heterogeneity. Ecological Modelling 93: 125-137.
15.  With, K. A., and T. O. Crist. 1995. Critical thresholds in species' responses to landscape structure. Ecology 76: 2446-2459.  Recipient of the 1996 Award for Best Paper published in the field of landscape ecology (US-IALE)
14.  Wiens, J. A., T. O. Crist, K. A. With, and B. T. Milne. 1995. Fractal patterns of insect movement in microlandscape mosaics. Ecology 76: 663-666.
13.  With, K. A. 1994. Ontogenetic shifts in how grasshoppers interact with landscape structure: an analysis of movement patterns. Functional Ecology 8: 477-485.
12.  With, K. A. 1994. Using fractal analysis to assess how species perceive landscape structure. Landscape Ecology 9: 25-36.
11.  With, K. A. 1994. The hazards of nesting near shrubs for a grassland bird, the McCown's Longspur. Condor 96: 1009-1019.   
10.  With, K. A. and D. R. Webb. 1993. Microclimate of ground nests: the relative importance of radiative cover and wind breaks for three grassland species. Condor 95: 401-413.   
9.  With, K. A., and M. L. Morrison. 1990. Flock formation of two parids in relation to cyclical seed production in a pinyon-juniper woodland. Auk 107: 522-532.   
8.  With, K. A., and R. P. Balda. 1990. Intersexual variation and factors affecting parental care in Western Bluebirds: a comparison of nestling and fledgling periods. Canadian Journal of Zoology 68: 733-742.   
7.  Morrison, M. L., K. A. With, I. C. Timossi, and K. A. Milne. 1987. Composition and temporal variation of flocks in the Sierra Nevada. Condor 89: 739-745.
6.  Block, W. M., K. A. With, and M. L. Morrison. 1987. On measuring bird habitat: influence of observer variability and sample size. Condor 89: 241-251.
5.  Morrison, M. L., K. A. With, I. C. Timossi, W. M. Block, and K. A. Milne. 1987. Foraging behavior of bark-foraging birds in the Sierra Nevada. Condor 89: 201-204.
4.  Morrison, M. L., and K. A. With. 1987. Interseasonal and intersexual resource partitioning in Hairy and White-headed woodpeckers. Auk 104: 225-233.
3.  Morrison, M. L., I. C. Timossi, and K. A. With. 1987. Development and testing of linear regression models predicting bird-habitat relationships. Journal of Wildlife Management 51: 247-253.
2.  Morrison, M. L., K. A. With, and I. C. Timossi. 1986. The structure of a forest bird community during winter and summer. Wilson Bulletin 98: 214-230.
1.  Morrison, M. L., I. C. Timossi, K. A. With, and P. N. Manley. 1985. Use of tree species by forest birds during winter and summer. Journal of Wildlife Management 49: 1098-1102.



Laboratory for Landscape and Conservation Ecology
2 Bushnell Hall
Division of Biology
Kansas State University
Manhattan, KS  66505  USA

Phone:  1-785-532-5040